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Section B. General Characters and Character States:
IX. Patterns
[A. Symmetry] [B. Arrangement Systems] [C. Branching Patterns]
- Actinomorphic or Radial. With floral parts radiate from center like spokes on wheel.
- Asymmetric. Without regularity in any dimension.
- Dorsiventral. Planate and having distinct dorsal and ventral surfaces, the two usually different.
- Equilateral. With halves or sides equal in shape and size.
- Inequilateral. With halves or sides unequal in shape and size.
- Irregular. With floral parts within a whorl dissimilar in shape and/ or size.
- Regular. With floral parts within a whorl similar in shape and size.
- Spherical. With multi-dimensional radial symmetry.
- Zygomorphic or Bilateral. With floral parts in two symmetrical halves.
- Anthotaxis. Arrangement of sporophylls, primarily reproductive in function.
- Carpotaxis. Arrangement of fruits, reproductive in function.
- Phyllotaxis. Arrangement of leaves, primarily photosynthetic in function.
- Rhizotaxis. Arrangement of roots.
- Semataxis. Arrangement of semaphylls (petals, sepals, tepals), primarily advertising (pollinator attracting) in function.
- Fibonacci Phyllotaxis. A fundamental type of leaf arrangement expressed as a fraction in which each succeeding fraction is the sum of the two previous numerators and the sum of the two previous denominators, i.e., 1/2, 1/3, 2/5, 3/8, 5/13, 8/21, etc. The numerator represents the number of turns or spirals around a stem before one leaf is directly above another and the denominator represents the number of leaves in the turns or spirals before one is directly above the other. 2/5 phyllotaxy would mean two twists and five leaves before one leaf is directly above the other or an angle of divergence of 144 between succeeding leaves in the stem (2/5 of 360 ). According to Leppik anthotaxis and semataxis do not necessarily follow the same pattern, with anthotaxis in Michelia cited as being in 2/7, 3/7, 3/8, and 4/10 systems of arrangement.
- Dichotomous. Branching into two equal parts.
- Flabellate. Fan-shaped branching.
- Monopodial. Branching with a main axis and reduced or missing laterals; excurrent.
- Without lateral branches. Main axis ending in a cluster of large leaves; e.g., Carica papaya.
- With continuous lateral branches. Main axis beset with laterals; e.g., Trema lamarckiana.
- With continuous but self-pruning lateral branches (excurrent). Main axis with upper laterals present and lower having fallen off; e.g., Pinus.
- With short-lived laterals. Main axis beset with phyllomorphic branches, appearing as compound leaves that may be deciduous; e.g., some tropical Rubiaceae, Pycnanthus.
- With spiral or whorled main laterals on main axis and distichous branches; e.g., Pycnanthus.
- With spiral or whorled main laterals and dichotomous branches; e.g., Phyllanthus.
- With tiered or clustered lateral branches. Main axis with periodic lateral branch production; e.g., Bombax.
- Sympodial. Branching without a main axis but with many, more or less, equal laterals.
- Leader displacement. Successive laterals take over leader growth producing umbrella-shaped trees; e.g., Acacia, Melia.; deliquescent.
- Leader displacement by two equal laterals (dichotomous substitution). Production of a forked branching system; e.g., Manihot.
- Leader displacement as a short-shoot (apposition). Usually a single lateral becomes dominant; e.g., Terminalia.
Note: Other types of stem growth and branching patterns occur, particularly in tropical woody plants but no adequate classification has been devised.
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